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    https://www2.mrc-lmb.cam.ac.uk/groups/JYL/PDF/e21600-download.pdf
    19 Dec 2016: For correspondence: jyl@mrc-. lmb.cam.ac.uk. Competing interests: The. authors declare that no. competing interests exist. Funding: See page 15. Received: 17 September 2016. Accepted: 14 November 2016. Published: 17 November 2016. Reviewing editor:

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